Breeding ecology of the Collared Flycatcher in forest-steppe deciduous forests of Sumy region. - N.P. Knysh. - Berkut. 12 (1-2). 2003. - P. 100-111.
The study was carried out on the station “Vakalivshchina” (51.01 N, 34.55 E) near Sumy mainly in 1999–2002. In middle-aged and old deciduous forest population density of the species remains at a level in 20–50 pairs/km2, in areas with many nest boxes it increases till 426 pairs/km2. In spring first Collared Flycatchers arrive from 1.04 (1979) till 3.05 (1987), on average 17.04 ±1,8 (standard error) days (n = 19). 535 cases of nesting were observed. 44 (8,2 %) nests were found in tree hollows, 478 (91,8 %) – in nest boxes for small birds. The earliest nests with first egg were recorded from 27.04 (2000) till 10.05 (1991, 1999), on average 5.05 ± 1,2 days (n = 13); the latest ones – 26.05 (1997, 1999) – 12.06 (1974), on average 3.06 ± 1,7 days (n = 10). The maximum (30,6 %) of clutches arrives during the third pentad of May (Table 1). Average date of beginning of egg laying (including repeated clutches) is 14.05 ± 0,4 days (n = 382, SD = 7,7). Average duration of phases of successful reproductive cycle, days: nest building – 4 (n = 4), pause before egg laying – 2,6 ± 0,2 (n = 47), egg laying – 6,47 ± 0,05 (n = 363), brooding – 13,2 ± 0,2 (n = 25), feeding of nestlings – 15,0 ± 0,2 (n = 25). Full clutches have 4–9, on average 6,47 ± 0,05 eggs (n = 363). Differences of average clutch sizes in separate years are described (Table 3). Number of eggs decreases during the breeding season (Table 4). Egg measurements depend on clutch size (Table 5), average values: 18,06 ± 0,02 x 13,36 ± 0,01 mm (15,6–21,0 x 11,4–14,8 mm; n = 1545). Their differences in separate years were studied (Table 6). Broods have on average 5,68 ± 0,10 (n = 193) hatchlings and 5,25 ± 0,13 (n = 162) fledglings (Table 7). Hatching productivity makes up on average 4,66 ± 0,19 (n = 183), fledging productivity – 4,01 ± 0,20 individuals per active nest (Table 8) or 74,3 and 63,9 % from eggs laid. Breeding success differs on years a little, but it depends on clutch size (Table 9). The largest losses of offspring (Table 10) are during brooding (20,9 %) and feeding of nestlings (10,4 %), the least ones – during egg laying (4,8 %). Predation (mainly by Dryomys nitedula) is the main reason of death (46,4 % of reproductive losses). [Russian].
Key words: Collared Flycatcher, Ficedula albicollis, Sumy region, ecology, breeding, clutch, breeding success, migration.
Adress: N.P. Knysh, Sumy Pedagogical University, Dep. of Zoology, Romenska str. 87, 40002, Sumy, Ukraine.

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